It's odd how natural laws tend to work the same way in Pokemon. For those of you who don't know, the Competitive Exclusion Principle states that no two organisms can occupy the same niche in an environment for an indefinite period of time; one of the two organisms will be driven to extinction. If you think for a second, you'll realize that this is true in Pokemon as well. Take, say, Gallade for example. Lucario and Gallade are, all in all, similar Pokemon in terms of the role that they play on teams. If you have a Lucario on your team, chances are that you don't also run a Gallade since the roles they generally play (aka not the Double Status set or whatever) are remarkably similar. Over time, when both of these Pokemon were commonplace in OU, people started to realize that Lucario's greater Speed, movepool, and typing enabled it to be the better sweeper. Hence, natural selection and the Competitive Exclusion Principle took effect, forcing Gallade out of the niche and into extinction (BL/UU). The same is true of Rotom-A and Dusknoir. Both are used as bulky Ghosts and spin blockers, but Rotom-A's better typing, ability, and movepool pushed it past its spectral competitor. When was the last time you saw a Dusknoir?
Dragons are probably the best example of natural selection and the CEP. Garchomp and Flygon originally occupied the same niche. Comparing them side by side, Garchamp clearly had the advantage over Flygon in stats and movepool. As such, Garchomp forced Flygon out of the niche and became the sole occupant. However, once Garchomp was removed from the environment, Flygon started to rise in usage and has recently flourished. You'll notice that Flygon and Salamence both thrive because they occupy different niches; Salamence is a DDer or a Mixed wall breaker while Flygon preforms revenge killing duties as a Scarfer. If both tried to only fulfill the position of a mixed wallbreaker, Salamence would most likely drive Flygon to 'extinction' due to its superior movepool and Special Attack. Salamence was about to push Dragonite into obscurity because they both filled essentially the same niche in OU, but then Dragonite got adaptations in the forms of Superpower and Extremespeed and it claimed a niche, however nebulous, for its own.
As Darwin said, it is not the strongest nor the most intelligent organism who survives, but the one who most efficiently adapts to a changing environment. This is why Pokemon such as Salamence, Gyarados, and Heatran have withstood the test of time and remained top players in OU. It doesn't matter how the metagame is structured at any given time because these Pokemon have the means to adapt to it. Pokemon such as Electivire have declined in usage because the changing metagame offers too much competition for it to fulfill its fundamental niche and results in it acquiring a much lower realized niche.
In nature, organisms often adapt to accommodate the adaptations of other organisms. This is true in Pokemon as well. The most common example of this that I can provide is the Scizor/Gengar relationship. Scizor's newest adaptation, Bullet Punch, provided it with the means to easily eliminate Gengar. Because of this, Gengar acquired a new set in the form of MysticGar. It was only because of the challenge presented by Scizor that the MysticGar set is viable. In any other situation, it would be perceived as a waste of a moveslot. Gengar's response to this change in the metagame allowed it to continue to function, and not go extinct as Pokemon such as the inflexible Porygon-Z and Electivire.
I'm not really sure where I'm going with this, but I think that it's interesting that natural laws govern the environment of competitive Pokemon. It's slightly unnerving to think that such aspects of nature have this sort of impact in something that is entirely virtual. Each teir's metagame is like its own ecosystem: they are constantly changing with cycles of different Pokemon filling specific niches as the metagame mutates. Nature seems to have the final say in everything, as everything that exists seems to fit into one pattern or another.
Dragons are probably the best example of natural selection and the CEP. Garchomp and Flygon originally occupied the same niche. Comparing them side by side, Garchamp clearly had the advantage over Flygon in stats and movepool. As such, Garchomp forced Flygon out of the niche and became the sole occupant. However, once Garchomp was removed from the environment, Flygon started to rise in usage and has recently flourished. You'll notice that Flygon and Salamence both thrive because they occupy different niches; Salamence is a DDer or a Mixed wall breaker while Flygon preforms revenge killing duties as a Scarfer. If both tried to only fulfill the position of a mixed wallbreaker, Salamence would most likely drive Flygon to 'extinction' due to its superior movepool and Special Attack. Salamence was about to push Dragonite into obscurity because they both filled essentially the same niche in OU, but then Dragonite got adaptations in the forms of Superpower and Extremespeed and it claimed a niche, however nebulous, for its own.
As Darwin said, it is not the strongest nor the most intelligent organism who survives, but the one who most efficiently adapts to a changing environment. This is why Pokemon such as Salamence, Gyarados, and Heatran have withstood the test of time and remained top players in OU. It doesn't matter how the metagame is structured at any given time because these Pokemon have the means to adapt to it. Pokemon such as Electivire have declined in usage because the changing metagame offers too much competition for it to fulfill its fundamental niche and results in it acquiring a much lower realized niche.
In nature, organisms often adapt to accommodate the adaptations of other organisms. This is true in Pokemon as well. The most common example of this that I can provide is the Scizor/Gengar relationship. Scizor's newest adaptation, Bullet Punch, provided it with the means to easily eliminate Gengar. Because of this, Gengar acquired a new set in the form of MysticGar. It was only because of the challenge presented by Scizor that the MysticGar set is viable. In any other situation, it would be perceived as a waste of a moveslot. Gengar's response to this change in the metagame allowed it to continue to function, and not go extinct as Pokemon such as the inflexible Porygon-Z and Electivire.
I'm not really sure where I'm going with this, but I think that it's interesting that natural laws govern the environment of competitive Pokemon. It's slightly unnerving to think that such aspects of nature have this sort of impact in something that is entirely virtual. Each teir's metagame is like its own ecosystem: they are constantly changing with cycles of different Pokemon filling specific niches as the metagame mutates. Nature seems to have the final say in everything, as everything that exists seems to fit into one pattern or another.