Ditto retaining the goofy smile reminds me of Meowth and Purrloin becoming bipedal by default, or even more akin to that particular Meowth gaining the ability to speak. Not sure if the bipedalism is a thing among Cat Pokemon, though.
Well Team Rocket's Meowth speaking human has remained a unique thing to him, however you're right that it's been retcon that walking on their hind legs is something the Meowth species can just do (and seemingly prefer to do over walking bipedal) even though TR's Meowth struggled to do so (and hindered him). And that's not even mentioning it's a default thing for the Regional Variants (and Meowth's Gigantamax). In comparison they do show Purrloin on all fours thanks to its default art (like if you look through the TCG art you'll sometimes see Purrloin quad while Meowth always biped after the early sets).
As for other cat Pokemon, let's see...
- Sneasel (if you count it as a cat) look completely biped.
- At first Skitty looks completely quad, but it does look like at times Skitty can stand on its hind legs as May's Skitty sometimes did using the Move Assist in the anime. That said, it could just be a momentary rearing up rather than learning how to stand balanced let alone walk on its hind legs.
- Glameow is an interesting case. Initially, while I thought it possibly could stand due to having similar proportions to Purrloin, I thought it has only ever been depicted as quad. BUT Bulbapedia just so happened to have two images that shows it somewhat bipedal. First is from the Adventures Manga, Johanna is doing stretch exercises and her Glameow is following along. Second isn't as concrete, it's from the anime, during one of Zoey's battles her Glameow goes into a bipedal pose while in midair (otherwise when on the ground it's quad).
- Trubbish looks completely biped (what? It was for some reason included in an image of cat Pokemon).
- Espurr looks completely biped.
- Litten & Sprigatito are quad, but doesn't stay like that for long to people's annoyance with quad-to-biped final Starter evos.
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